Praeratavites daohugou Rasnitsyn, Ansorge & Zhang, gen. et sp. n., Grimmaratavites mirabilis Rasnitsyn, Ansorge & Zhang, gen. et sp. n., and Praeparyssites orientalis Rasnitsyn, Ansorge & Zhang, sp. n. are described from the Lower Jurassic (Lower Toarcian) of Grimmen, NE Germany (G. mirabilis) and Middle Jurassic of Daohugou. Karatavitidae Rasnitsyn, 1963, is transferred from Ephialtitoidea into now paraphyletic infraorder Orussomorpha, and elevated to superfamily Karatavitoidea. The described genera, jointly with those earlier described in Karatavitidae and Paroryssidae, are found to form a smooth succession that model evolutionary transition from the basalmost Vespina toward the extant Orussidae.