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(Scarabaeidae: Scarabaeinae) forms an important part of the dung beetle fauna in much of the Western Hemisphere. Here a phylogeny for
, including 49
sp., and 12 outgroup taxa, is proposed. Parsimony analysis of 67 morphological characters, and one biogeographical character produced 629 equally parsimonious trees of 276 steps.
, the putative sister taxon is nested well within the subgenus
, implying that
might ultimately need to be synonymized with
. Species groups of Edmonds (1994) recovered as monophyletic are
. An 'unscaled' equal weighting analysis yielded 57,149 equally parsimonious trees of 372 steps. The strict consensus of these trees yielded a monophyletic
with the inclusion of
. Bootstrap values are relatively low and some clades are unresolved.
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