An updated key to the millipede-hunting subgenus Aganhyboma Kolbe, 1893 of the genus Deltochilum Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae), with description of a new species from Bolivia and Peru

in Insect Systematics & Evolution
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Deltochilum (Aganhyboma) mariafernandae sp. nov., from Santa Cruz, Bolivia and Cuzco, Peru is described based on differences in external and male genital morphology. Its diagnostic characters and an identification key to the species of the subgenus are provided.

An updated key to the millipede-hunting subgenus Aganhyboma Kolbe, 1893 of the genus Deltochilum Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae), with description of a new species from Bolivia and Peru

in Insect Systematics & Evolution

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References

Cano E.B. (1998) Deltochilum valgum acropyge Bates (Coleoptera: Scarabaeidae: Scarabaeinae): Habits and Distribution. Coleopterists Bulletin 52: 174178.

González-Alvarado A. & Vaz-de-Mello F. (2014) Taxonomic review of the subgenus Hybomidium Shipp, 1897 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). Annales de la Société entomologique de France (N.S.): International Journal of Entomology 50: 431476.

Halffter G. (2003) Tribu Scarabaeini. In: Morón M.Á. (ed.) Atlas de los escarabajos de México. Coleoptera: Lamellicornia. Familias Scarabaeidae Trogidae Passalidae y Lucanidae Argania editio Barcelona: pp. 2143.

Larsen T.H. Lopera A. Forsyth A. & Génier F. (2009) From coprophagy to predation: a dung beetle that kills millipedes. Biology Letters 5: 152155.

Pereira F.S. & Martínez A. (1956) Os gêneros de Canthonini americanos (Col. Scarabaeidae). Revista Brasileira de Entomologia 6: 91192.

Schoolmeesters P. (2017) Scarabs: World Scarabaeidae Database. In: Roskov Y. Abucay L. Orrell T. et al. (eds.) Species 2000 & ITIS Catalogue of Life. Available from URL: www.catalogueoflife.org/col. (updated 27 Febr 2017; accessed 17 Apr 2017).

Silva F.A.B. Louzada J. & Vaz-de-Mello F. (2015) A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 3925: 451504.

Silva F.A.B. Vidaurre T. Vaz-de-Mello F. & Louzada J. (2012) Predatory behaviour in Deltochilum: convergent evolution or a primitive character within a clade?. Journal of Natural History 46: 13591367.

Tarasov S.I. & Génier F. (2015) Innovative bayesian and parsimony phylogeny of dung beetle (Coleoptera, Scarabaeidae, Scarabaeinae) enhanced by ontology-based partitioning of morphological characters. Plos one 10: 186.

Tarasov S.I. & Solodovnikov A.Y. (2011) Phylogenetic analyses reveal reliable morphological markers to classify mega-diversity in Onthophagini dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae). Cladistics 27: 490528.

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    Deltochilum (Aganhyboma) mariafernandae sp. nov.. 1. Dorsal view. 2. Head. 3. Lateral view of elytra (black arrow shows the basal carina of seventh interstria; white arrow shows the basal carina of ninth interstria; red arrow shows the pseudepipleural carina). Figure 4. Hypomera of Deltochilum (A.) schefflerorum. Figure 5. Hypomera of Deltochilum (A.) arturoi. Figure 6. Hypomera of Deltochilum (A.) mariafernandae sp. nov.. Figure 7. Hypomera of Deltochilum (A.) acanthus.

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    Deltochilum (Aganhyboma) mariafernandae sp. nov.. 8. Detail of the third elytral interstria. 9. Detail of the first elytral stria. 10. Apical elytral tubercles (arrow shows apical tubercle of second interstria). Figure 11. Apical elytral tubercles in Deltochilum (A.) schefflerorum (arrow shows the apex of second interstria); Figures 12–19. Aedeagus and endophallic sclerites in Deltochilum (A.) mariafernandae sp. nov. 12. Parameres (dorsal view). 13. Parameres (ventral view). 14. Complex of axial and subaxial sclerites (A + SA). 15. Fronto-lateral peripheral sclerite (FLP). 16. Superior right peripheral sclerite (SRP). 17. Aedeagus (lateral view). 18. Submedial lobe of endophallus. 19. Lamella copulatrix sclerites (LC).

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    Known distribution of acropyge species complex.

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