The limits of the Palaearctic Region, like those of other biogeographical regions, are arbitrarily defined (Map 1). For practical reasons, the boundaries of the Palaearctic Region usually follow national boundaries. The region includes Europe, Africa north of the Sahara, and Asia except for the part that is arbitrarily defined as belonging to the Oriental Region.

For each species and subspecies, an outline of its present distribution is given using symbols. The Palaearctic Region is further subdivided into three main parts, Europe (letter E), North Africa (letter N) and Asia (letter A).

Europe includes the Azores, Iceland and Turkey west of the Bosporus. The eastern boundaries are a matter of controversy. In the present Catalogue, Europe includes Russia west of the main range of the Ural Mountains (Permsky Krai, Bashkortostan Republic and Orenburg Oblast entirely, in Sverdlovsk Oblast and Chelyabinsk Oblast only the parts west of 60ºE) and a small part of Kazakhstan west of the Ural River. The south-eastern boundaries are the northern political boundaries of Georgia, Azerbaijan and the Caspian and Black seas. The limits of Ukraine reflect the position in 2021; Luhansk Oblast, Donetsk Oblast, Zaporizhzhia Oblast and Kherson Oblast are parts of Ukraine. Crimea is listed separately, mainly due to its unique fauna. Abkhazia is for practical reasons given under Georgia.

In the previous Edition the abbreviation YU designated Serbia and Montenegro (at that time constituting the Federal Republic of Yugoslavia) only, however, many authors adopted it for records from entire former Yugoslavia. During the preparation phase of the Catalogue Serbia and Montenegro were split, what was not reflected in the published version. Subsequently also Kosovo was declared as independent therefore in the present edition each of the countries of former Yugoslavia has its own abbreviation to avoid any further confusions and misapplications.

North Africa includes Morocco (incl. Western Sahara), Algeria, Tunisia, Libya, Egypt west of the Suez Canal, and the Canary, Madeira and Salvage islands.

Asia includes Sinai and the Arabian Peninsula (including Socotra), Turkey east of the Bosporus, the Caucasian republics (Georgia, Armenia and Azerbaijan), the Middle East and Central Asian countries, Russia east of the main range of the Ural Mountains, Korea, Japan (including the Ryukyu Archipelago and the Japanese Pacific Islands), the entire People’s Republic of China, Taiwan, Bhutan, Nepal, North India along the base of the Himalayas (Arunachal Pradesh, Uttarakhand (north-western area of former Uttar Pradesh), Himachal Pradesh, Jammu and Kashmir) and all of Pakistan. Thus, India is the only state for which strict political boundaries were not followed.

Two-letter symbols provided for countries are arranged in alphabetical order within each subdivision. The two-letter symbols for major areas of Russia, North Indian states, and three-letter symbols for provinces of mainland China and for Taiwan (see Map 3) are placed in brackets after the respective country code. Russia is subdivided into seven major sub-regions (Map 2). In previous editions of the Palaearctic Catalogue Siberia was divided into West Siberia and East Siberia only. Now, Siberia is divided into three units: East Siberia, South Siberia and West Siberia, which aim to better describe its zoogeographical diversity. South Siberia includes the entire republics of Altai, Tuva and Khakassia, Kemerovo Region without the part north of Kemerovo, the extreme south of Altai Province, the southern part of Krasnoyarsk Province to the latitude of Krasnoyarsk, the South-West of Irkutsk Region and the western part of the Republic of Buryatia to Selenga River in the east. The symbols RU for Russia and CH for the People’s Republic of China, without specification in parentheses, are used only in cases where precise information was not available.

Manchuria (also Mandzhuria or Manchukuo) was a geographical region in Northeast Asia often referred to by various authors, in particular by G. G. Jacobson, until 1949. It is presently divided between China (Heilongjiang, Jilin and Liaoning and eastern part of Inner Mongolia) and Russia (south-eastern part of Russian Far East including Sakhalin); e.g. https://en.wikipedia.org/wiki/Manchuria. In many recent publications, old Manchurian records from the Chinese part were automatically considered to be from Heilongjiang (compare, for example, the record of Parnops atriceps Pic, Jacobson (1910b) and its interpretation by Li et al. (2008) through the citation in Tan et al. (2005)). However, at least since 1905, Jacobson also had material from Jilin and Liaoning, and many of his Manchurian records thus belong to these provinces, which can be verified by checking locality labels under respective specimens. The following localities are situated in Jilin (collector Serikov): “Цайцзягоу, ю. Харбина” (Tsaitzyagou, S of Harbin) [= Caijiagou, 45°06′55″N, 126°08′29″E]; and Liaoning (collector Bortkevich, mostly in surroundings of “Мамайкая” (Mamaikaya)) [= Mamajecun, 41°25′36″N 122°51′25″E]): “Чендятунь, з. Мамайкая, Мугден” (Chendyatun’, W of Mamaikaya, Mukden) [= Zhengjiatun, 41°28′38″N 122°45′10″E], “Худяпуза, з. Мамайкая, Мугден” (Hudyapuza, W of Mamaikaya, Mukden) [= Houjia Gangzi, 41°24′09″N, 122°47′34″E], “Сышагоу, ю.з. Мамайкая, Мугден” (Syshagou, SW of Mamaikaya, Mukden) [approximately 41°21′N 122°46′E], “Лангаши” (Langashi) [this locality was not found but is certainly situated in the same area]. We thank Alexey Moseyko for providing this essential information about Manchuria.

Some, usually older, distributional records cannot be accommodated within the structure of the Catalogue (e.g., Arabia, Caucasus, North India, Siberia). Such information is given in quotation marks (e.g., “Caucasus”) after the last symbol of the subdivision. Distributional records of species-group taxa with isolated occurrence, either endemic to very small geographic area (e.g. islands, mountains etc.) or widely distributed in one area but restricted in another area, may be specified in parentheses.

The extralimital distribution of Palaearctic species is indicated by three letter symbols in boldface located at the end of the respective geographical information. The extralimital regions for the purposes of the Catalogue are defined as follows (see Map 1): Nearctic (NAR): north of Mexico; Neotropical (NTR): south of the United States; Afrotropical (AFR): south of the North African states included in the Palaearctic Region; Oriental (ORR): areas south of the People’s Republic of China and Taiwan, areas south of Himalayan India, the Philippines, Malaysia and Indonesia west of the Lydekker line; Australian (AUS): south and east of the Lydekker line, and Pacific (PAC). Introductions are indicated by the letter “i” (e.g., Ei: GB).

In general, the presented distributional information is based on published records scattered in an enormous number of publications. Therefore, the second-level geographic information is not necessarily exhaustive, it should rather be considered as a baseline for future faunal research. The authors of some groups in the current catalogue were unable to find published sources or voucher specimens for some countries or areas listed in the previous Edition. These distribution data are included in the present catalogue but also listed in the Comments. In addition, to present distributional information as accurate as possible, new faunistic records are presented in this Edition, in New Acts chapter separately for each subfamily or tribe.

Map 1
Map 1

The limits of the geographic Regions as defined for the purpose of this Catalogue

Map 2
Map 3
Map 3

Subdivisions and provinces of the People’s Republic of China, and Taiwan

Geographical Symbols

E

Europe

AL

Albania

AN

Andorra

AU

Austria

AZ

Azores

BE

Belgium

BH

Bosnia Herzegovina

BU

Bulgaria

BY

Belarus

CR

Croatia

CZ

Czech Republic

DE

Denmark

EN

Estonia

FA

Faeroe Islands

FI

Finland

FR

France (incl. Corsica, Monaco)

GB

Great Britain (incl. Channel Islands)

GE

Germany

GR

Greece (incl. Crete)

HU

Hungary

IC

Iceland

IR

Ireland

IT

Italy (incl. Sardinia, Sicily, San Marino)

KO

Kosovo

KR

Crimea

KZ

Kazakhstan

LA

Latvia

LS

Liechtenstein

LT

Lithuania

LU

Luxembourg

MA

Malta

MD

Moldavia

ME

Montenegro

NL

The Netherlands

NM

North Macedonia

NR

Norway

PL

Poland

PT

Portugal

RO

Romania

RU

Russia

SB

Serbia

SK

Slovakia

SL

Slovenia

SP

Spain (incl. Gibraltar)

SV

Sweden

SZ

Switzerland

TR

Turkey

UK

Ukraine

N

North Africa

AG

Algeria

CI

Canary Islands

EG

Egypt

LB

Libya

MO

Morocco (incl. Western Sahara)

MR

Madeira Archipelago

SG

Salvage Islands

TU

Tunisia

A

Asia

AB

Azerbaijan

AE

United Arab Emirates

AF

Afghanistan

AR

Armenia

BA

Bahrain

BT

Bhutan

CH

China

CY

Cyprus

GG

Georgia

ID

India

IN

Iran

IQ

Iraq

IS

Israel (incl. Palestine)

JA

Japan

JO

Jordan

KI

Kyrgyzstan

KU

Kuwait

KZ

Kazakhstan

LE

Lebanon

MG

Mongolia

NC

North Korea

NP

Nepal

OM

Oman

PA

Pakistan

QA

Qatar

RU

Russia

SA

Saudi Arabia

SC

South Korea

SI

Egypt: Sinai

SY

Syria

TD

Tajikistan

TM

Turkmenistan

TR

Turkey

UZ

Uzbekistan

YE

Yemen (incl. Socotra)

Russia: European part

CT

Central European Territory

NT

North European Territory

ST

South European Territory

Russia: Asian part

ES

East Siberia

FE

Far East

SS

South Siberia

WS

West Siberia

China: Provinces, Autonomous Regions or Municipalities, and Taiwan

ANH

Anhui (Anhwei)

BEI

Beijing (Peking or Peiping)

CHQ

Chongqing

FUJ

Fujian (Fukien)

GAN

Gansu (Kansu)

GUA

Guandong (Kwantung)

GUI

Guizhou (Kweichow)

GUX

Guangxi (Kwangsi)

HAI

Hainan

HEB

Hebei (Hopeh)

HEI

Heilongjiang (Heilungkiang)

HEN

Henan (Honan)

HKG

Hong Kong

HUB

Hubei (Hupeh)

HUN

Hunan

JIA

Jiangsu (Kiangsu)

JIL

Jilin (Kirin)

JIX

Jiangxi (Kiangsi)

LIA

Liaoning

MAC

Macao

NIN

Ningxia (Ningsia)

NMO

Nei Mongol (Inner Mongolia)

QIN

Qinghai (Tsinghai)

SCH

Sichuan (Szechwan)

SHA

Shaanxi (Shensi)

SHG

Shanghai

SHN

Shandong (Shantung)

SHX

Shanxi (Shansi)

TAI

Taiwan (Formosa)

TIA

Tianjin (Tsiensin)

XIN

Xinjiang (Sinkiang)

XIZ

Xizang (Tibet)

YUN

Yunnan

ZHE

Zhejiang (Chekiang)

India: Provinces

AP

Arunachal Pradesh

HP

Himachal Pradesh

KA

Jammu and Kashmir, Ladakh

SD

Sikkim, Darjeeling District

UT

Uttarakhand (Uttaranchal, former part of Uttar Pradesh)

World Zoogeographical Regions

AFR

Afrotropical Region

AUS

Australian Region

NAR

Nearctic Region

NTR

Neotropical Region

ORR

Oriental Region

PAC

Pacific Region

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