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Julia Ostner and Oliver Schülke

1. Introduction Two decades ago primatologists compiled what was then known about primate male affiliative relationships (e.g., van Hooff & van Schaik, 1992, 1994; Hill & van Hooff, 1994 ; Strier, 1994a). It was proposed that affiliation and cooperation evolved less frequently in males

Carel P. Van Schaik and Jan A.R.A.M. Van Hooff

MALE BONDS: AFILLIATIVE RELATIONSHIPS AMONG NONHUMAN PRIMATE MALES by JAN A.R.A.M. VAN HOOFF1) and CAREL P. VAN SCHAIK2) (1 Ethologie & Socio-oecologie Groep, Universiteit Utrecht, Utrecht, the Netherlands; 2 Biological Anthropology & Anatomy, Duke University, Durham NC, U.S.A.) (With 2 Figures

Serge Wich and Elisabeth Sterck

, Behavioural Biology, P.O. Box 80086, 3508 TB, Utrecht, The Netherlands; 4 Ethology Research, Animal Science Department, Biomedical Primate Research Centre, Postbus 3306, 2280 GH Rijswijk, The Netherlands) (Accepted: 22 June 2007) Summary During between-group encounters in primates, males often defend mates

Erin P. Riley, Cristina Sagnotti, Monica Carosi and Ngakan Putu Oka

macaques . — Int. J. Primatol. 13 : 97 - 105 . Preuschoft S. Paul A. ( 2000 ). Dominance, egalitarianism, and stalemate: an experimental approach to male–male competition in Barbary macaques . — In: Primate males ( Kappeler P.M. , ed.). Cambridge University Press

Jeannette P. Hanby and Conrad E. Brown

inhibit intromission delay the development of the young primate male's ability to distinguish correctly between postures, partners, and situations. The copulatory patterns of females seem less affected by unsatisfactory rearing condi- tions, but the sociosexual patterns of contact may be disturbed. 3

Katharine M. Jack and Erin P. Riley

articles focusing on affiliative relationships among primate males was published by this journal (Behaviour 130 (1994): 143–337). This Special Issue brought together a series of 10 papers (including a preface by Hill & van Hooff), three of which focused on Neotropical primates, two on the genus Macaca

Denise N. Gabriel, Lisa Gould and Elizabeth A. Kelley

over access to receptive females (Wrangham, 1980 ; van Hooff & van Schaik, 1994; van Schaik, 1996). Thus, early studies of interactions between primate males have focused largely on aggressive and dominance behaviours (e.g., Kaufman, 1967 ; Packer, 1979 ; Colvin, 1983 ; Janson, 1986 ; Smuts

Sterck, Becher, Wich and Assink

-over (Wee- den & Falls, 1959; Emlen, 1971; Ydenberg et al ., 1988; Whiting, 1999). Also in primates males may be recognised by their call alone, because in- dividual differences in primate calls have been shown for a variety of species ( e.g . chimpanzees, Pan troglodytes : Marler & Hobett, 1975; vervet

Julie A. Teichroeb, Eva C. Wikberg, Nelson Ting and Pascale Sicotte

animals ( Harcourt A.H. de Waal F.B.M. , eds). Oxford University Press , Oxford , p.  415 - 444 . Cords M. ( 2000 ). The number of males in guenon groups . — In: Primate males: causes and consequences of variation in group composition ( Kappeler P.M. , ed.). Cambridge

Christopher Young, Oliver Schülke and Julia Ostner

of within-group male coalitions seems limited, because female-based models assume philopatric allies and competition over a sharable resource, whereas most primate males are dispersing and compete in a zero-sum game over fertilizations. Other models of primate male coalition formation examined payoff