Orthoclads from Eocene Amber from Sakhalin (Diptera: Chironomidae, Orthocladiinae)

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Four Middle Eocene orthoclad species, Heterotrissocladius naibuchi sp. n., Paraphaenocladius nadezhdae sp. n., Pseudosmittia kodrulae sp. n. and Smittia sukachevae sp. n.. are described and figured. Based on the combination of the diptera fauna composition and paleobotanical data, suggestions on the climate and habitats of the Sakhalin amber forest are given.

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